拟髭蟾属

拟髭蟾属
	Leptobrachium hasseltii（英语：Leptobrachium hasseltii）
科学分类
界：	动物界 Animalia
门：	脊索动物门 Chordata
纲：	两栖纲 Amphibia
目：	无尾目 Anura
科：	角蟾科 Megophryidae
属：	拟髭蟾属 Leptobrachium; Tschudi（英语：Johann Jakob von Tschudi）, 1838
模式种
	Leptobrachium hasseltii（英语：Leptobrachium hasseltii）; Tschudi（英语：Johann Jakob von Tschudi）, 1838
物种
	约35种，详见内文
异名
	髭蟾属 Vibrissaphora Liu, 1945;

拟髭蟾属（学名：Leptobrachium）是无尾目角蟾科的一属动物，原髭蟾属（Vibrissaphora）已并入该属^[1]^[2]。

分布

本属物种分布于华南、印度东北、东南亚、巽他陆棚上的岛屿，以及菲律宾^[3]。它们的特点是有结实的身体和细长的后肢^[4]。

形态特征

本属物种眼睛虹膜的颜色是分辨各物种的关键：例如波普拟髭蟾（波普拟髭蟾）就有一双蓝眼睛；有些物种的虹色单一，但也有物种的虹蟆只有上半圈有颜色，而下半圈只有深色^[5]。

分类

演化树

拟髭蟾属的姐妹群、跟拟髭蟾属位处同一演化支的包括有齿突蟾属及齿蟾属^[3]^[7]。

亚属

本属已确认的亚属有以下两个^[3]：

同名亚属Leptobrachium，以及
Vibrissaphora，即过往的髭蟾属，模式种是峨眉髭蟾（Vibrissaphora boringii）。

这两者的差异，就只是在形态上它们的雄性动物的上唇在繁殖季节有没有长出婚刺（英语：nuptial pad）。后来的遗传分析不支持过往这种基于形态的分离，但仍然表明存在两个不同的演化支，所以把两支演化支视为同属的亚属比较合适。从生物地理学来说，两者在地理上也有不同的分布：Vibrissaphora亚属分布于华南及中南半岛，而指名亚属分布于马来半岛及华莱士线西北方的马来群岛^[5]。除此以外，两者并没有其他在形态学上可分辨的特征^[8]。

物种

截至2019年^[update]，Frost的文献认为本属现时有36个获确认的物种^[3]，以及一个有待确认的物种：

Leptobrachium abbotti (Cochran, 1926)
哀牢髭蟾 Leptobrachium ailaonicum (Yang, Chen, and Ma, 1983)
Leptobrachium banae Lathrop, Murphy, Orlov & Ho, 1998
波普拟髭蟾 Leptobrachium bompu Sondhi & Orlov, 2011^[9]）
峨眉髭蟾 Leptobrachium boringii (Liu, 1945)）
Leptobrachium buchardi Ohler, Teynié & David, 2004^[10]
沙巴拟髭蟾 Leptobrachium chapaense (Bourret, 1937)）
广西拟髭蟾 Leptobrachium guangxiense Fei, Mo, Ye & Jiang, 2009） Leptobrachium gunungense
Leptobrachium gunungense Malkmus, 1996
海南拟髭蟾 Leptobrachium hainanensis Ye & Fei In Ye, Fei & Hu, 1993）
Leptobrachium hasseltii Tschudi, 1838
Leptobrachium hendricksoni Taylor, 1962
华深拟髭蟾 Leptobrachium huashen Fei and Ye, 2005）
Leptobrachium ingeri Hamidy, Matsui, Nishikawa, and Belabut, 2012
Leptobrachium kanowitense Hamidy, Matsui, Nishikawa, and Belabut, 2012
Leptobrachium kantonishikawai Hamidy and Matsui, 2014^[11]
雷山髭蟾 Leptobrachium leishanense (Liu and Hu, 1973)）
Leptobrachium leucops Stuart, Rowley, Tran, Le & Hoang, 2011^[12]
崇安髭蟾 Leptobrachium liui (Pope, 1947)）
Leptobrachium lumadorum Brown, Siler, Diesmos & Alcala, 2010
Leptobrachium lunatum Stuart, Som, Neang, Hoang, Le, Dau, Potter & Rowley, 2020^[13]
Leptobrachium mangyanorum Brown, Siler, Diesmos & Alcala, 2010
Leptobrachium masatakasatoi Matsui, 2013^[8]
Leptobrachium montanum Fischer, 1885
Leptobrachium mouhoti Stuart, Sok & Neang, 2006
Leptobrachium ngoclinhense Orlov, 2005
Leptobrachium nigrops Berry & Hendrickson, 1963
原髭蟾 Leptobrachium promustache (Rao, Wilkinson, and Zhang, 2006)）
Leptobrachium pullum (Smith, 1921)
Leptobrachium rakhinensis Wogan, 2012^[14]
Leptobrachium smithi Matsui, Nabhitabhata & Panha, 1999^[15]
Leptobrachium sylheticum Al-Razi, Maria, and Poyarkov, 2021^[16]
Leptobrachium tagbanorum Brown, Siler, Diesmos & Alcala, 2010
丹那沙林拟髭蟾 Leptobrachium tenasserimense Pawangkhanant, Poyarkov, Duong, Naiduangchan, and Suwannapoom, 2018，泰国西部丹那沙林^[17]
腾冲拟髭蟾 Leptobrachium tengchongense Yang, Wang, & Chan, 2016）^[18]
Leptobrachium waysepuntiense Hamidy & Matsui, 2010^[19]
Leptobrachium xanthops Stuart, Phimmachak, Seateun & Sivongxay, 2012^[20]
Leptobrachium xanthospilum Lathrop, Murphy, Orlov & Ho, 1998

参考文献

^ Yuan, Zhiyong; Chen, Jinmin; Wu, Yunhe; Li, Xianqi; Che, Jing. Revision of the list of amphibian species in Yunnan Province. Biodiversity Science. 2022-04-20, 30 (4): 21470. doi:10.17520/biods.2021470.
^ Vibrissaphora Liu, 1945. GBIF. [2022-07-13]. （原始内容存档于2022-07-13）（英语）.
^ ^3.0 ^3.1 ^3.2 ^3.3 Frost, Darrel R. Leptobrachium Tschudi, 1838. Amphibian Species of the World: an Online Reference. Version 6.0. American Museum of Natural History. 2017 [2017-06-25]. （原始内容存档于2019-04-06）（英语）.
^ Matsui, Masafumi; Jarujin Nabhitabhata; Somsak Panha. On Leptobrachium from Thailand with a description of a new species (Anura: Pelobatidae). Japanese Journal of Herpetology. 1999, 18 (1): 19–29. doi:10.5358/hsj1972.18.1_19 .
^ ^5.0 ^5.1 Matsui, M.; Hamidy, A.; Murphy, R. W.; Khonsue, W.; Yambun, P.; Shimada, T.; Ahmad, N.; Belabut, D. M.; Jiang, J. P. Phylogenetic relationships of megophryid frogs of the genus Leptobrachium (Amphibia, Anura) as revealed by mtDNA gene sequences. Molecular Phylogenetics and Evolution. 2010, 56 (1): 259–272. PMID 20302957. doi:10.1016/j.ympev.2010.03.014.
^ Hudson, C. M.; Fu, J. Male-biased sexual size dimorphism, resource defense polygyny, and multiple paternity in the Emei moustache toad (Leptobrachium boringii). PLOS ONE. 2013, 8 (6): e67502. Bibcode:2013PLoSO...867502H. PMC 3696078 . PMID 23840725. doi:10.1371/journal.pone.0067502 .
^ Pyron, A. R.; Wiens, J. J. A large-scale phylogeny of Amphibia including over 2800 species, and a revised classification of extant frogs, salamanders, and caecilians. Molecular Phylogenetics and Evolution. 2011, 61 (2): 543–583. PMID 21723399. doi:10.1016/j.ympev.2011.06.012.
^ ^8.0 ^8.1 Matsui, M. A new Leptobrachium (Vibrissaphora) from Laos (Anura: Megophryidae). Current Herpetology. 2013, 32 (2): 182–189. S2CID 85577993. doi:10.5358/hsj.32.182. hdl:2433/216851 .
^ Sondhi, S.; Orlov, A. A blue-eyed Leptobrachium (Anura: Megophryidae) from Arunachal Pradesh, India (PDF). Zootaxa. 2011, 2912: 28–36 [2021-10-24]. doi:10.11646/zootaxa.2912.1.2. （原始内容存档 (PDF)于2021-10-27）.
^ Ohler, A.; Teynié, A. & David, P. A green-eyed Leptobrachium (Anura: Megophryidae) from southern Laos (PDF). Raffles Bulletin of Zoology（英语：Raffles Bulletin of Zoology）. 204, 52 (2): 695–700. （原始内容 (PDF)存档于2012-03-16）.
^ Hamidy A, Matsui M 2014. A New Species of Leptobrachium from the Kelabit Highland, Northwestern Borneo (Anura, Megophryidae). Curr Herpetol. 33(1): 57-67.
^ Stuart, B. L.; Rowley, J. J.; Tran, Dao T. A.; Le, Duong T. T. & Hoang, H. D. The Leptobrachium (Anura: Megophryidae) of the Langbian Plateau, southern Vietnam, with description of a new species (PDF). Zootaxa. 2011, 2804: 25–40 [2021-10-24]. doi:10.11646/zootaxa.2804.1.3 . （原始内容存档 (PDF)于2019-07-24）.
^ Bryan L. Stuart, Hannah E. Som, Thy Neang, Huy Duc Hoang, Duong Thi Thuy Le, Vinh Quang Dau, Kathy Potter and Jodi J. L. Rowley. 2020. Integrative Taxonomic Analysis reveals A New Species of Leptobrachium (Anura: Megophryidae) from north-eastern Cambodia and central Vietnam. Journal of Natural History. DOI: 10.1080/00222933.2020.1756498
^ Wogan, G.O.U. 2012. A New Species of Leptobrachium from Myanmar (Anura: Megophryidae). Zootaxa. 3415: 23-36.
^ Dipankar Dutta, Abhijit Das, Amalesh Dutta, Jayanta Gogoi and Saibal Sengupta. 2013. Taxonomic Status and Distribution of Leptobrachium smithi Matsui, Nabhitabhata & Panha, 1999 (Anura: Megophryidae) in India with New Locality Records. Tropical Natural History. 13(2): 87-95
^ Leptobrachium sylheticum Al-Razi, Maria, and Poyarkov, 2021 | Amphibian Species of the World. [2021-10-24]. （原始内容存档于2021-10-26）.
^ Parinya Pawangkhanant, Nikolay A. Poyarkov, Tang Van Duong, Mali Naiduangchan and Chatmongkon Suwannapoom. 2018. A New Species of Leptobrachium (Anura, Megophryidae) from western Thailand. PeerJ. 6:e5584. DOI: 10.7717/peerj.5584
^ Jian-Huan Yang, Ying-Yong Wang and Bosco Pui-Lok Chan. 2016. A New Species of the Genus Leptobrachium (Anura: Megophryidae) from the Gaoligongshan Mountain Range, China. Zootaxa. 4150(2); DOI: 10.11646/zootaxa.4150.2.3
^ Hamidy, A.; Matsui, M. A new species of blue-eyed Leptobrachium (Anura: Megophryidae) from Sumatra, Indonesia (PDF). Zootaxa. 2010, 2395: 34–44 [2021-10-24]. doi:10.11646/zootaxa.2395.1.3. （原始内容存档 (PDF)于2016-03-03）（英语）.
^ Stuart, B. L.; Phimmachak, S.; Seateun, S.; Sivongxay, N. A new Leptobrachium (Anura: Megophryidae) from the highlands of southeastern Laos (PDF). Zootaxa. 2012, 2155: 29–37 [2021-10-24]. （原始内容存档 (PDF)于2019-07-24）.

外部链接

[1] Yuan, Zhiyong; Chen, Jinmin; Wu, Yunhe; Li, Xianqi; Che, Jing. Revision of the list of amphibian species in Yunnan Province. Biodiversity Science. 2022-04-20, 30 (4): 21470. doi:10.17520/biods.2021470.

[2] Vibrissaphora Liu, 1945. GBIF. [2022-07-13]. （原始内容存档于2022-07-13）（英语）.

[Frost-3] 3.0 ^3.1 ^3.2 ^3.3 Frost, Darrel R. Leptobrachium Tschudi, 1838. Amphibian Species of the World: an Online Reference. Version 6.0. American Museum of Natural History. 2017 [2017-06-25]. （原始内容存档于2019-04-06）（英语）.

[Matsui-4] Matsui, Masafumi; Jarujin Nabhitabhata; Somsak Panha. On Leptobrachium from Thailand with a description of a new species (Anura: Pelobatidae). Japanese Journal of Herpetology. 1999, 18 (1): 19–29. doi:10.5358/hsj1972.18.1_19 .

[Matsui2010-5] 5.0 ^5.1 Matsui, M.; Hamidy, A.; Murphy, R. W.; Khonsue, W.; Yambun, P.; Shimada, T.; Ahmad, N.; Belabut, D. M.; Jiang, J. P. Phylogenetic relationships of megophryid frogs of the genus Leptobrachium (Amphibia, Anura) as revealed by mtDNA gene sequences. Molecular Phylogenetics and Evolution. 2010, 56 (1): 259–272. PMID 20302957. doi:10.1016/j.ympev.2010.03.014.

[HudsonFu-6] Hudson, C. M.; Fu, J. Male-biased sexual size dimorphism, resource defense polygyny, and multiple paternity in the Emei moustache toad (Leptobrachium boringii). PLOS ONE. 2013, 8 (6): e67502. Bibcode:2013PLoSO...867502H. PMC 3696078 . PMID 23840725. doi:10.1371/journal.pone.0067502 .

[7] Pyron, A. R.; Wiens, J. J. A large-scale phylogeny of Amphibia including over 2800 species, and a revised classification of extant frogs, salamanders, and caecilians. Molecular Phylogenetics and Evolution. 2011, 61 (2): 543–583. PMID 21723399. doi:10.1016/j.ympev.2011.06.012.

[Matsui2013-8] 8.0 ^8.1 Matsui, M. A new Leptobrachium (Vibrissaphora) from Laos (Anura: Megophryidae). Current Herpetology. 2013, 32 (2): 182–189. S2CID 85577993. doi:10.5358/hsj.32.182. hdl:2433/216851 .

[9] Sondhi, S.; Orlov, A. A blue-eyed Leptobrachium (Anura: Megophryidae) from Arunachal Pradesh, India (PDF). Zootaxa. 2011, 2912: 28–36 [2021-10-24]. doi:10.11646/zootaxa.2912.1.2. （原始内容存档 (PDF)于2021-10-27）.

[10] Ohler, A.; Teynié, A. & David, P. A green-eyed Leptobrachium (Anura: Megophryidae) from southern Laos (PDF). Raffles Bulletin of Zoology（英语：Raffles Bulletin of Zoology）. 204, 52 (2): 695–700. （原始内容 (PDF)存档于2012-03-16）.

[11] Hamidy A, Matsui M 2014. A New Species of Leptobrachium from the Kelabit Highland, Northwestern Borneo (Anura, Megophryidae). Curr Herpetol. 33(1): 57-67.

[12] Stuart, B. L.; Rowley, J. J.; Tran, Dao T. A.; Le, Duong T. T. & Hoang, H. D. The Leptobrachium (Anura: Megophryidae) of the Langbian Plateau, southern Vietnam, with description of a new species (PDF). Zootaxa. 2011, 2804: 25–40 [2021-10-24]. doi:10.11646/zootaxa.2804.1.3 . （原始内容存档 (PDF)于2019-07-24）.

[13] Bryan L. Stuart, Hannah E. Som, Thy Neang, Huy Duc Hoang, Duong Thi Thuy Le, Vinh Quang Dau, Kathy Potter and Jodi J. L. Rowley. 2020. Integrative Taxonomic Analysis reveals A New Species of Leptobrachium (Anura: Megophryidae) from north-eastern Cambodia and central Vietnam. Journal of Natural History. DOI: 10.1080/00222933.2020.1756498

[14] Wogan, G.O.U. 2012. A New Species of Leptobrachium from Myanmar (Anura: Megophryidae). Zootaxa. 3415: 23-36.

[15] Dipankar Dutta, Abhijit Das, Amalesh Dutta, Jayanta Gogoi and Saibal Sengupta. 2013. Taxonomic Status and Distribution of Leptobrachium smithi Matsui, Nabhitabhata & Panha, 1999 (Anura: Megophryidae) in India with New Locality Records. Tropical Natural History. 13(2): 87-95

[16] Leptobrachium sylheticum Al-Razi, Maria, and Poyarkov, 2021 | Amphibian Species of the World. [2021-10-24]. （原始内容存档于2021-10-26）.

[17] Parinya Pawangkhanant, Nikolay A. Poyarkov, Tang Van Duong, Mali Naiduangchan and Chatmongkon Suwannapoom. 2018. A New Species of Leptobrachium (Anura, Megophryidae) from western Thailand. PeerJ. 6:e5584. DOI: 10.7717/peerj.5584

[18] Jian-Huan Yang, Ying-Yong Wang and Bosco Pui-Lok Chan. 2016. A New Species of the Genus Leptobrachium (Anura: Megophryidae) from the Gaoligongshan Mountain Range, China. Zootaxa. 4150(2); DOI: 10.11646/zootaxa.4150.2.3

[19] Hamidy, A.; Matsui, M. A new species of blue-eyed Leptobrachium (Anura: Megophryidae) from Sumatra, Indonesia (PDF). Zootaxa. 2010, 2395: 34–44 [2021-10-24]. doi:10.11646/zootaxa.2395.1.3. （原始内容存档 (PDF)于2016-03-03）（英语）.

[20] Stuart, B. L.; Phimmachak, S.; Seateun, S.; Sivongxay, N. A new Leptobrachium (Anura: Megophryidae) from the highlands of southeastern Laos (PDF). Zootaxa. 2012, 2155: 29–37 [2021-10-24]. （原始内容存档 (PDF)于2019-07-24）.

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查论编按亚目分类的现存的无尾目
界动物界Animalia · 门脊索动物门 Chordata · 亚门脊椎动物亚门 Chordata · 总纲四足总纲 Tetrapoda · 纲两栖纲 Amphibia
始蛙亚目 Archaeobatrachia	尾蟾科 Ascaphidae · 铃蟾科 Bombinatoridae · 盘舌蟾科 Discoglossidae · 滑跖蟾科 Leiopelmatidae
中蛙亚目 Mesobatrachia	角蟾科 Megophryidae · 锄足蟾科 Pelobatidae · 潜蟾科 Pelodytidae · 负子蟾科Pipidae · 北美锄足蟾科 Scaphiopodidae · 异舌蟾科 Rhinophrynidae
新蛙亚目 Neobatrachia	真齿蛙科 Amphignathodontidae · 隐林蛙科 Aromobatidae · 节蛙科 Arthroleptidae · 短头蟾科 Brachycephalidae · 蟾蜍科 Bufonidae · 瞻星蛙科 Centrolenidae · 角花蟾科 Ceratophryidae · Craugastoridae · 丛蛙科 Dendrobatidae · 沼蟾科 Heleophrynidae · 扩角蛙科 Hemiphractidae · 肩蛙科 Hemisotidae · 雨蛙科 Hylidae · 芦蛙科 Hyperoliidae · 细趾蟾科 Leptodactylidae · 曼蛙科 Mantellidae · 姬蛙科 Microhylidae · 龟蟾科 Myobatrachidae · 箱头蛙科 Pyxicephalidae · 蛙科 Ranidae · 树蛙科 Rhacophoridae · 尖吻达蛙科 Rhinodermatidae · 塞舌蛙科 Sooglossidae